The study on the origin, evolution and phylogeny of fungi was originally based on comparative morphological and cytological data. After 1980s, with the development of science and technology and the wide application of new technologies, such as the study of G-C content, the composition and structure of cell wall polysaccharide, the pattern of enzyme precipitation in tryptophan biosynthesis pathway of various fungi, two different synthesis pathways of lysine and the study of rRNA sequence, etc. And promoted the study on the origin and evolution of fungi. It is not clear how long fungi have existed on the earth, and there is no definite conclusion about the origin of fungi. Fungi are similar to plants in some characteristics, but similar to animals in some aspects. From 1980s to 1990s, according to the comparative study of nutrition patterns, fungi are neither plants nor animals, but an independent biological group-Fungus Kingdom. Diversity of origin: According to the morphology of sexual organs and mating methods, fungi are considered to come from algae. Hyphaceae evolved from Prorocentrum; Aquatic fungi evolved from algae-free; Mucor evolved from synbiotic algae; Ascomycetes and basidiomycetes evolved from red algae. These algae changed from autotrophic to heterotrophic because of the loss of pigment, and the physiological changes caused morphological changes. This is a pluralistic view of the origin of fungi. The origin theory of flagellates: It is believed that most fungi originated from a primitive aquatic organism-flagellates, single cells, with one or several flagella, some with pigments such as chlorophyll, some without pigments, and evolved into pigmented algae and pigments-free fungi. Both fungi and algae originated from flagellate organisms. Lifestyle: aquatic fungi are primitive, and the evolution process is from aquatic to terrestrial. It is speculated that they may return to aquatic habits during the evolution process. Therefore, it is considered that the flagellated swimming spores are primitive and the immobile static spores are relatively evolved. Nutrition: Saprophytic lifestyle is primitive lifestyle, and parasitic lifestyle is more advanced than saprophytic lifestyle. The obligate parasitic lifestyle is more advanced than the facultative parasitic lifestyle, and the highest lifestyle is the specific obligate parasitic lifestyle. Fungal structure: from simple to complex, and then from complex to degradation, the special structure is lost, thus simplifying the structure. The wide application of new technology provides a scientific basis for correcting the origin and evolution of fungi. Basidiomycetes.